Ethics
Appearance
This is strikingly parallel to modern accounts of cooperation in evolutionary game theory, where agents that recognize their fitness as coupled to the fitness of others outcompete purely self-regarding agents.
The bridge between these evolutionary dynamics and individual moral cognition is moral psychology, which reveals that moral emotions — guilt, shame, gratitude — function as commitment devices calibrated by social interaction. An ethics that ignores how moral cognition actually works is designing for agents that do not exist.